Tuning is important but especially if it's a carb'd application, air speed and booster signal at lower RPMs are crucial. They are directly related to how responsive an engine is under light throttle. Also when larger amounts of valve overlap come into play the tuning of the exhaust can make a huge difference in torque below the peak RPM. Lots of tests showing how collector extensions on open headers give a massive improvement in torque and the longer they get the lower the RPMs where torque is increased.