what crankshaft could i use?

perfacar said
the 273 cranks were fitted to 904 trans. the poly headed 318 were fitted to 727 trans. therefore, the back register is different between the two. find a machinist, have him bore the back register out to 727 size. it's done in a lathe. I know, I do them.
How is it then that I have a '69 273 out of a Fury that had a 904 that is now in an '85 pickup with a 727?